Epipactis (abbr.: Epcts.) is not exactly the most sexy of all terrestrial orchids. It occurs in Europe, Northern America, Africa and Asia in forests, grasslands and in the middle of cities. A number of interesting hybrids have hit the market in recent years, increasing attention for this genus.
The name Epipactis derives from a classical name used by Theophrastus (circa 350 B.C.) for a plant used to curdle milk. About 25 species found worldwide, mostly in Europe (15 species) and Asia. Two species are found in the U.S. - E. gigantea and E. helleborine. The latter is thought to have arrived in North America from Europe in the 19th century and grows in the northeastern U.S.
Epipactis species are normally grown in the garden, they appear to have a strong relation with their fungus, but this depends strongly from species to species.
Plants don't have a tuber but thick roots that radiate from the shoot.
Epipactis species are typically grown in the garden, some can be started in potculture but will soon move to the backyard as they multiply. Most of them are winterhard down to -15°C and can withstand Summer maxima of 35°C. Hybrids often need protection in winter - normally if one of the parents is Mediterranean. Most species can take full sunlight, even at noon, but this is not required. Conditions that are too shady are reflected in a sparse number of flowers. Epipactis species seem to like a medium that keeps the roots humid and they can take a lot of rain in winter. There are three rules for mixing a soil: neutral pH (5.5 - 7.0), good drainage (max 5% clay) and not too many organic ingredients (max 25%). You can use regular commercial gardenmix compost as a base, add grit/sand/perlite/vermiculite/pumice/gravel for drainage and loam/peat/fen soil/sphagnum/coco peat/oyster shells to adjust the pH. Water with rainwater.
Repot when fully dormant but take care in not braking roots. We normally leave the roots and shoot in a pocket of original soil and plant this in the new medium. You can fertilise the plants, preferable with 10-20% of the advised quantity.
Epipactis gigantea (notes on culture by Nina Rach)
Virtually always found in wet places, commonly known as the "stream orchid," "giant helleborine," "chatterbox," and "false lady's slipper." It's the most common native orchid in California and ranges to west Texas, northern Mexico, and southern Canada. There are usually 10 or more green, ovate-lanceolate leaves per plant, alternating on the stem, each from 5-11 cm wide and up to 25 cm long. There is a variety native to California that has wine-red leaves. In cultivation, the plants usually reach 30-50 cm tall. The foliage is deciduous - dies back in the fall. After the foliage turns brown, the new shoots begin to spread away from the rhizome.
Plants bloom to about 1 meter high, with 12-20 flowers per inflorescence. The flowers are 4 to 4.5 cm across (up to 5.5 cm), and usually face the same direction. Sepals are usually dark green, petals rose with dark veining. The (usually red) lip is deeply three-lobed, hinged in the middle, with yellow lateral lobes. The epichile is elongated and vibrates in slight breezes (hence, the "chatterbox" orchid) and the ovary is straight. Flowers have four pollinia. White flower forms and peloric forms are known. Blooming season is early March to early October; the species blooms in 39 months from seed.
Habitat: Found from sea level to 2600 m, always with a constant source of water at the roots, including wet cliff-faces, salty beaches, road-cuts, mountain bogs, hot springs, and sandbars in streams. The rhizomes are slender.
Culture: Mix: peat moss, sand, perlite, some charcoal, humus. You may also add powdered lime and bonemeal. The colony of plants at Kew Gardens is grown beside a small stream in a rock garden, in peat, loam, and leaf mold compost in partial shade, along with dactylorhizas and Skunk Cabbage. A colony is also growing at the Geneva Botanic Garden in a rock garden (drier and sunnier than Kew) along with Cypripedium reginae. Keep in medium-shade.
Best to repot just as plants are breaking dormancy; however, Carson Whitlow recommends handling plants in the autumn, just before dormancy. Tip of leading growth bud should be level with top of compost.
Keep pot in a saucer or trough of water, in bright light. Can grow outdoors in Houston year-round; might benefit if refrigerated or kept cool in some way over the winter. The plants will not tolerate a hard freeze. Suitable for the Holman and Whitlow artificial bog techniques (see Cribb & Bailes), or moist rock gardens.
Epipactis palustris
The Marsh helleborine likes it a bit wetter than most orchids. Don't let the medium dry out but avoid stagnant water, the soil should be water retentive and rich in lime. Forms dense clumps after a number of years because it multiplies vegetatively and flowers end of April/begin May. Seeds are difficult to germinate in culture, they need to take up water and then by stratified for a period of time. This species is suited for potculture and gardenculture, mostly in the formed of raised beds. Some growers plant them around ponds. They normally triple in number every year when grown in pots. Soil should be alkaline to neutral, acidic soils will not kill the plants but they don't seem to prefer it. Young plants are more sensitive to the pH than adult plants. You can use general garden compost with a generous serving of vermiculite, perlite, sand and grit for drainage. Add crushed oystershell or limestone to increase pH. Mix some Sphagnum moss in the bottom of the pot to serve as a wick. Place in a low saucer of water, the soil should not become sogging wet (use a pot that is high enough). Top of with half a centimeter of grit to prevent the moist soil from attracting insects and to slow down evaporation. Water from the top every now and then to aerate the medium.
Epipactis gigantea
Syrphid fly
A large population of Epipactis atropurpureum on 40-year-old zinc wastes in Chrzanow, southern Poland, where the soil contained high concentrations of cadmium, lead and zinc, was studied for heavy metal content and mycorrhizal development. Rhizomes of the orchid contained extremely high levels of these heavy metals, and the copper content was five times higher then that found in the soil. Heavy metal contents in rhizomes were 10 times (zinc) to 100 times (lead, cadmium) higher than in the shoots. These results suggested the accumulation and biofiltering of metals within rhizomes. But how? E. atropurpureum almost always has mycorrhizal roots: the fungus penetrates the rhizome forming complex hyphal coils. Most of the toxic elements had accumulated in the fungal coils in the rhizomes: lead, iron, zinc, calcium, sulphur and aluminium were in much higher concentrations in the fungal coils than in the surrounding cells. Mycorrhizal fungi may play an important role in heavy metal sequestration and detoxification, allowing the plant to survive in extremely polluted places. 1)
Epcts. pallustris is sold occasionally in gardencentres.
Ground-Orchids.de sells a number of hybrids
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec | Location | |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Epipactis microphylla | Belgium | ||||||||||||
| Epipactis palustris | U.K. | ||||||||||||
| Epipactis atrorubens | North Wales, Scotland, Ireland | ||||||||||||
| Epipactis purpurata | South-Central and South-East England, U.K | ||||||||||||
| Epipactis helleborine | Wales, Centr. Scotland, England, U.K | ||||||||||||
| Epipactis leptochila | U.K | ||||||||||||
| Epipactis dunensis | Central U.K | ||||||||||||
| Epipactis sancta | Lindisfarne National Nature Reserve, U.K | ||||||||||||
| Epipactis phyllanthes | Central Southern England, U.K |
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec | Location | |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Epipactis africana | 1984 | Uganda | |||||||||||
| Epipactis atrorubens | 1953 | 1932 | Austria | ||||||||||
| Epipactis atrorubens | 2007 | 1982 | 1973 | France | |||||||||
| Epipactis atrorubens | 2001 | 1934 | Germany | ||||||||||
| Epipactis atrorubens | 1929 | Greece | |||||||||||
| Epipactis atrorubens | 1974 | 1973 | Italy | ||||||||||
| Epipactis atrorubens | 1995 | Poland | |||||||||||
| Epipactis atrorubens | 1974 | Slovenia | |||||||||||
| Epipactis atrorubens | 1930 | 1936 | 1957 | Spain | |||||||||
| Epipactis atrorubens | 2002 | 1994 | 1994 | 1987 | Switzerland | ||||||||
| Epipactis condensata | 1963 | 1948 | Cyprus | ||||||||||
| Epipactis condensata | 1986 | 1948 | 1978 | Turkey | |||||||||
| Epipactis densifolia | 1886 | Switzerland | |||||||||||
| Epipactis dunensis | 1991 | Great Britain | |||||||||||
| Epipactis gigantea | 1935 | 1951 | Afghanistan | ||||||||||
| Epipactis gigantea | 1973 | 1973 | Pakistan | ||||||||||
| Epipactis gigantea | 1934 | 1902 | U.S.A. | ||||||||||
| Epipactis helleborine | 1968 | Austria | |||||||||||
| Epipactis helleborine | 1935 | 2007 | France | ||||||||||
| Epipactis helleborine | 1992 | 1982 | 1985 | Germany | |||||||||
| Epipactis helleborine | 1930 | Great Britain | |||||||||||
| Epipactis helleborine | 1951 | Greece | |||||||||||
| Epipactis helleborine | 1983 | 1983 | India | ||||||||||
| Epipactis helleborine | 1977 | 1974 | 1974 | Iran | |||||||||
| Epipactis helleborine | 2000 | 2004 | Italy | ||||||||||
| Epipactis helleborine | 1937 | Morocco | |||||||||||
| Epipactis helleborine | 1930 | Netherlands | |||||||||||
| Epipactis helleborine | 1982 | 1973 | Pakistan | ||||||||||
| Epipactis helleborine | 1957 | Spain | |||||||||||
| Epipactis helleborine | 1968 | 1974 | 1991 | 1968 | Switzerland | ||||||||
| Epipactis helleborine | 1982 | 1982 | 1989 | 1972 | Turkey | ||||||||
| Epipactis helleborine subsp. helleborine | 1907 | Austria | |||||||||||
| Epipactis helleborine subsp. helleborine | 1962 | 1950 | France | ||||||||||
| Epipactis helleborine subsp. helleborine | 1929 | Germany | |||||||||||
| Epipactis helleborine subsp. helleborine | 1930 | 1929 | Greece | ||||||||||
| Epipactis helleborine subsp. helleborine | 1936 | Spain | |||||||||||
| Epipactis helleborine subsp. helleborine | 1952 | 1958 | 1953 | Switzerland | |||||||||
| Epipactis helleborine subsp. helleborine | 1951 | Turkey | |||||||||||
| Epipactis helleborine subsp. orbicularis | 1937 | Greece | |||||||||||
| Epipactis helleborine subsp. orbicularis | 1981 | 1998 | Switzerland | ||||||||||
| Epipactis helleborine subsp. tremolsii | 2003 | 2001 | Spain | ||||||||||
| Epipactis kleinii | 2003 | Spain | |||||||||||
| Epipactis latifolia | 1950 | Germany | |||||||||||
| Epipactis latifolia | 1950 | 1950 | Great Britain | ||||||||||
| Epipactis latifolia | 1927 | Greece | |||||||||||
| Epipactis latifolia | 1961 | India | |||||||||||
| Epipactis latifolia | 1954 | Netherlands | |||||||||||
| Epipactis latifolia | 1930 | 1953 | Switzerland | ||||||||||
| Epipactis latifolia | 1952 | 1950 | 1959 | Turkey | |||||||||
| Epipactis leptochila | 2007 | 2006 | France | ||||||||||
| Epipactis leptochila | 1950 | Great Britain | |||||||||||
| Epipactis leptochila | 1997 | 1997 | Switzerland | ||||||||||
| Epipactis leptochila subsp. neglecta | 2007 | France | |||||||||||
| Epipactis leptochila subsp. neglecta | 2002 | Great Britain | |||||||||||
| Epipactis leptochila subsp. neglecta | 2000 | Switzerland | |||||||||||
| Epipactis mairei | 2004 | China | |||||||||||
| Epipactis microphylla | 1904 | 1890 | Austria | ||||||||||
| Epipactis microphylla | 2007 | 2007 | 1926 | France | |||||||||
| Epipactis microphylla | 1995 | Germany | |||||||||||
| Epipactis microphylla | 1953 | 1929 | Greece | ||||||||||
| Epipactis microphylla | 1975 | 1972 | Iran | ||||||||||
| Epipactis microphylla | 1954 | Italy | |||||||||||
| Epipactis microphylla | 1930 | 2003 | Spain | ||||||||||
| Epipactis microphylla | 1995 | 1978 | 1975 | 1944 | Switzerland | ||||||||
| Epipactis microphylla | 1987 | 1958 | Turkey | ||||||||||
| Epipactis muelleri | 1992 | 2007 | 2007 | France | |||||||||
| Epipactis muelleri | 1988 | Switzerland | |||||||||||
| Epipactis palustris | 1905 | Austria | |||||||||||
| Epipactis palustris | 2007 | 2007 | 1971 | 1964 | France | ||||||||
| Epipactis palustris | 2007 | 1935 | Germany | ||||||||||
| Epipactis palustris | 1929 | 1929 | Greece | ||||||||||
| Epipactis palustris | 1954 | Hungary | |||||||||||
| Epipactis palustris | 1970 | 1972 | Iran | ||||||||||
| Epipactis palustris | 1989 | 1997 | Italy | ||||||||||
| Epipactis palustris | 1957 | Spain | |||||||||||
| Epipactis palustris | 2002 | 1997 | 1984 | 1987 | Switzerland | ||||||||
| Epipactis palustris | 1951 | Turkey | |||||||||||
| Epipactis palustris f. ericetorum | 1950 | Great Britain | |||||||||||
| Epipactis persica | 1973 | Afghanistan | |||||||||||
| Epipactis persica | 2001 | Georgia | |||||||||||
| Epipactis persica | 1974 | 1974 | Iran | ||||||||||
| Epipactis persica | 1973 | 1973 | Pakistan | ||||||||||
| Epipactis persica | 1981 | 1978 | 1972 | 1913 | Turkey | ||||||||
| Epipactis persica subsp. moravica | 1997 | Germany | |||||||||||
| Epipactis phyllanthes var. phyllanthes | 1958 | France | |||||||||||
| Epipactis phyllanthes var. phyllanthes | 1951 | 1951 | Great Britain | ||||||||||
| Epipactis phyllanthes | 2002 | Great Britain | |||||||||||
| Epipactis phyllanthes | 1957 | Spain | |||||||||||
| Epipactis placentina | 1994 | Italy | |||||||||||
| Epipactis placentina | 1999 | Switzerland | |||||||||||
| Epipactis pontica | 1989 | Austria | |||||||||||
| Epipactis pontica | 1978 | Turkey | |||||||||||
| Epipactis rechingeri | 1974 | 1974 | Iran | ||||||||||
| Epipactis troodi | 1963 | Cyprus | |||||||||||
| Epipactis troodi | 1959 | Turkey | |||||||||||
| Epipactis veratrifolia | 1973 | 1967 | 1948 | Afghanistan | |||||||||
| Epipactis veratrifolia | 1994 | 1992 | 1949 | Cyprus | |||||||||
| Epipactis veratrifolia | 1949 | 1975 | 1974 | Iran | |||||||||
| Epipactis veratrifolia | 1982 | Pakistan | |||||||||||
| Epipactis veratrifolia | 1982 | 1951 | Turkey | ||||||||||
| Epipactis viridiflora subsp. kuenkeleana | 1996 | Azerbaydzhan | |||||||||||
| Epipactis viridiflora | 1989 | Bosnia-Herzegovina | |||||||||||
| Epipactis viridiflora | 1961 | Czech Republic | |||||||||||
| Epipactis viridiflora | 1963 | 2007 | 2007 | 2006 | 2006 | France | |||||||
| Epipactis viridiflora | 1996 | Germany | |||||||||||
| Epipactis viridiflora | 1966 | 1996 | 1970 | Switzerland |
Below is the list of names accepted by the Kew Monocot Checklist. For the complete list - including invalid names - please click here.
World Checklist for Epipactis. With the permission of the Trustees of the Royal Botanic Gardens, Kew Published on the Internet