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Culture & Care for the genus Epipactis

Introduction to the genus

Epipactis (abbr.: Epcts.) is not exactly the most sexy of all terrestrial orchids. It occurs in Europe, Northern America, Africa and Asia in forests, grasslands and in the middle of cities. A number of interesting hybrids have hit the market in recent years, increasing attention for this genus.
The name Epipactis derives from a classical name used by Theophrastus (circa 350 B.C.) for a plant used to curdle milk. About 25 species found worldwide, mostly in Europe (15 species) and Asia. Two species are found in the U.S. - E. gigantea and E. helleborine. The latter is thought to have arrived in North America from Europe in the 19th century and grows in the northeastern U.S.
Epipactis species are normally grown in the garden, they appear to have a strong relation with their fungus, but this depends strongly from species to species.
Plants don't have a tuber but thick roots that radiate from the shoot.


General cultivation techniques

Epipactis species are typically grown in the garden, some can be started in potculture but will soon move to the backyard as they multiply. Most of them are winterhard down to -15°C and can withstand Summer maxima of 35°C. Hybrids often need protection in winter - normally if one of the parents is Mediterranean. Most species can take full sunlight, even at noon, but this is not required. Conditions that are too shady are reflected in a sparse number of flowers. Epipactis species seem to like a medium that keeps the roots humid and they can take a lot of rain in winter. There are three rules for mixing a soil: neutral pH (5.5 - 7.0), good drainage (max 5% clay) and not too many organic ingredients (max 25%). You can use regular commercial gardenmix compost as a base, add grit/sand/perlite/vermiculite/pumice/gravel for drainage and loam/peat/fen soil/sphagnum/coco peat/oyster shells to adjust the pH. Water with rainwater.
Repot when fully dormant but take care in not braking roots. We normally leave the roots and shoot in a pocket of original soil and plant this in the new medium. You can fertilise the plants, preferable with 10-20% of the advised quantity.

Specific guidelines

Epipactis gigantea (notes on culture by Nina Rach)
Virtually always found in wet places, commonly known as the "stream orchid," "giant helleborine," "chatterbox," and "false lady's slipper." It's the most common native orchid in California and ranges to west Texas, northern Mexico, and southern Canada. There are usually 10 or more green, ovate-lanceolate leaves per plant, alternating on the stem, each from 5-11 cm wide and up to 25 cm long. There is a variety native to California that has wine-red leaves. In cultivation, the plants usually reach 30-50 cm tall. The foliage is deciduous - dies back in the fall. After the foliage turns brown, the new shoots begin to spread away from the rhizome.
Plants bloom to about 1 meter high, with 12-20 flowers per inflorescence. The flowers are 4 to 4.5 cm across (up to 5.5 cm), and usually face the same direction. Sepals are usually dark green, petals rose with dark veining. The (usually red) lip is deeply three-lobed, hinged in the middle, with yellow lateral lobes. The epichile is elongated and vibrates in slight breezes (hence, the "chatterbox" orchid) and the ovary is straight. Flowers have four pollinia. White flower forms and peloric forms are known. Blooming season is early March to early October; the species blooms in 39 months from seed.
Habitat: Found from sea level to 2600 m, always with a constant source of water at the roots, including wet cliff-faces, salty beaches, road-cuts, mountain bogs, hot springs, and sandbars in streams. The rhizomes are slender.
Culture: Mix: peat moss, sand, perlite, some charcoal, humus. You may also add powdered lime and bonemeal. The colony of plants at Kew Gardens is grown beside a small stream in a rock garden, in peat, loam, and leaf mold compost in partial shade, along with dactylorhizas and Skunk Cabbage. A colony is also growing at the Geneva Botanic Garden in a rock garden (drier and sunnier than Kew) along with Cypripedium reginae. Keep in medium-shade.
Best to repot just as plants are breaking dormancy; however, Carson Whitlow recommends handling plants in the autumn, just before dormancy. Tip of leading growth bud should be level with top of compost.
Keep pot in a saucer or trough of water, in bright light. Can grow outdoors in Houston year-round; might benefit if refrigerated or kept cool in some way over the winter. The plants will not tolerate a hard freeze. Suitable for the Holman and Whitlow artificial bog techniques (see Cribb & Bailes), or moist rock gardens.

Epipactis palustris
The Marsh helleborine likes it a bit wetter than most orchids. Don't let the medium dry out but avoid stagnant water, the soil should be water retentive and rich in lime. Forms dense clumps after a number of years because it multiplies vegetatively and flowers end of April/begin May. Seeds are difficult to germinate in culture, they need to take up water and then by stratified for a period of time. This species is suited for potculture and gardenculture, mostly in the formed of raised beds. Some growers plant them around ponds. They normally triple in number every year when grown in pots. Soil should be alkaline to neutral, acidic soils will not kill the plants but they don't seem to prefer it. Young plants are more sensitive to the pH than adult plants. You can use general garden compost with a generous serving of vermiculite, perlite, sand and grit for drainage. Add crushed oystershell or limestone to increase pH. Mix some Sphagnum moss in the bottom of the pot to serve as a wick. Place in a low saucer of water, the soil should not become sogging wet (use a pot that is high enough). Top of with half a centimeter of grit to prevent the moist soil from attracting insects and to slow down evaporation. Water from the top every now and then to aerate the medium.

In-vitro culture

Hybridisation notes

Pollinators

Epipactis gigantea
Syrphid fly

Pests and diseases

Trivia

A large population of Epipactis atropurpureum on 40-year-old zinc wastes in Chrzanow, southern Poland, where the soil contained high concentrations of cadmium, lead and zinc, was studied for heavy metal content and mycorrhizal development. Rhizomes of the orchid contained extremely high levels of these heavy metals, and the copper content was five times higher then that found in the soil. Heavy metal contents in rhizomes were 10 times (zinc) to 100 times (lead, cadmium) higher than in the shoots. These results suggested the accumulation and biofiltering of metals within rhizomes. But how? E. atropurpureum almost always has mycorrhizal roots: the fungus penetrates the rhizome forming complex hyphal coils. Most of the toxic elements had accumulated in the fungal coils in the rhizomes: lead, iron, zinc, calcium, sulphur and aluminium were in much higher concentrations in the fungal coils than in the surrounding cells. Mycorrhizal fungi may play an important role in heavy metal sequestration and detoxification, allowing the plant to survive in extremely polluted places. 1)

References

Commercial sources

Epcts. pallustris is sold occasionally in gardencentres.
Ground-Orchids.de sells a number of hybrids

Flowering calendar

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec Location
Epipactis microphylla            Belgium
Epipactis palustris            U.K.
Epipactis atrorubens            North Wales, Scotland, Ireland
Epipactis purpurata            South-Central and South-East England, U.K
Epipactis helleborine            Wales, Centr. Scotland, England, U.K
Epipactis leptochila            U.K
Epipactis dunensis            Central U.K
Epipactis sancta            Lindisfarne National Nature Reserve, U.K
Epipactis phyllanthes            Central Southern England, U.K

Detailed data: Aggregated observation dates from the GBIF.org network.
A colored month means an observation in habitat or collection date of a herbarium specimen with flower or inflorescence. The number refers to the year of last observation or collection.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec Location
Epipactis africana       1984     Uganda
Epipactis atrorubens       19531932    Austria
Epipactis atrorubens      20071982 1973   France
Epipactis atrorubens       20011934    Germany
Epipactis atrorubens       1929     Greece
Epipactis atrorubens       19741973    Italy
Epipactis atrorubens      1995      Poland
Epipactis atrorubens       1974     Slovenia
Epipactis atrorubens    19301936 1957     Spain
Epipactis atrorubens      2002199419941987   Switzerland
Epipactis condensata       19631948    Cyprus
Epipactis condensata    1986 19481978     Turkey
Epipactis densifolia        1886    Switzerland
Epipactis dunensis       1991     Great Britain
Epipactis gigantea      19351951     Afghanistan
Epipactis gigantea      19731973     Pakistan
Epipactis gigantea     1934 1902     U.S.A.
Epipactis helleborine        1968    Austria
Epipactis helleborine      19352007     France
Epipactis helleborine      199219821985    Germany
Epipactis helleborine        1930    Great Britain
Epipactis helleborine      1951      Greece
Epipactis helleborine       19831983    India
Epipactis helleborine      197719741974    Iran
Epipactis helleborine       20002004    Italy
Epipactis helleborine      1937      Morocco
Epipactis helleborine         1930   Netherlands
Epipactis helleborine       19821973    Pakistan
Epipactis helleborine       1957     Spain
Epipactis helleborine      1968197419911968   Switzerland
Epipactis helleborine     1982198219891972    Turkey
Epipactis helleborine subsp. helleborine       1907    Austria
Epipactis helleborine subsp. helleborine      19621950    France
Epipactis helleborine subsp. helleborine      1929     Germany
Epipactis helleborine subsp. helleborine     19301929     Greece
Epipactis helleborine subsp. helleborine    1936       Spain
Epipactis helleborine subsp. helleborine     195219581953    Switzerland
Epipactis helleborine subsp. helleborine      1951     Turkey
Epipactis helleborine subsp. orbicularis     1937      Greece
Epipactis helleborine subsp. orbicularis     19811998     Switzerland
Epipactis helleborine subsp. tremolsii     20032001     Spain
Epipactis kleinii      2003      Spain
Epipactis latifolia       1950     Germany
Epipactis latifolia       19501950    Great Britain
Epipactis latifolia     1927       Greece
Epipactis latifolia       1961     India
Epipactis latifolia        1954    Netherlands
Epipactis latifolia       19301953    Switzerland
Epipactis latifolia     195219501959     Turkey
Epipactis leptochila       2007 2006   France
Epipactis leptochila        1950    Great Britain
Epipactis leptochila       19971997    Switzerland
Epipactis leptochila subsp. neglecta      2007     France
Epipactis leptochila subsp. neglecta      2002     Great Britain
Epipactis leptochila subsp. neglecta      2000     Switzerland
Epipactis mairei      2004      China
Epipactis microphylla       19041890    Austria
Epipactis microphylla      20072007 1926   France
Epipactis microphylla       1995     Germany
Epipactis microphylla      19531929     Greece
Epipactis microphylla     1975  1972    Iran
Epipactis microphylla      1954      Italy
Epipactis microphylla    1930 2003      Spain
Epipactis microphylla      1995197819751944   Switzerland
Epipactis microphylla      19871958     Turkey
Epipactis muelleri      199220072007    France
Epipactis muelleri        1988    Switzerland
Epipactis palustris       1905     Austria
Epipactis palustris      2007200719711964   France
Epipactis palustris      20071935     Germany
Epipactis palustris       19291929    Greece
Epipactis palustris       1954     Hungary
Epipactis palustris      19701972     Iran
Epipactis palustris      19891997     Italy
Epipactis palustris       1957     Spain
Epipactis palustris      2002199719841987   Switzerland
Epipactis palustris       1951     Turkey
Epipactis palustris f. ericetorum      1950     Great Britain
Epipactis persica     1973       Afghanistan
Epipactis persica       2001     Georgia
Epipactis persica       19741974    Iran
Epipactis persica       19731973    Pakistan
Epipactis persica      1981197819721913   Turkey
Epipactis persica subsp. moravica       1997    Germany
Epipactis phyllanthes var. phyllanthes     1958      France
Epipactis phyllanthes var. phyllanthes      1951 1951   Great Britain
Epipactis phyllanthes       2002     Great Britain
Epipactis phyllanthes       1957     Spain
Epipactis placentina       1994     Italy
Epipactis placentina       1999     Switzerland
Epipactis pontica        1989    Austria
Epipactis pontica       1978     Turkey
Epipactis rechingeri       19741974    Iran
Epipactis troodi       1963     Cyprus
Epipactis troodi       1959     Turkey
Epipactis veratrifolia     197319671948     Afghanistan
Epipactis veratrifolia   19941992 1949      Cyprus
Epipactis veratrifolia    19491975 1974     Iran
Epipactis veratrifolia       1982     Pakistan
Epipactis veratrifolia      19821951     Turkey
Epipactis viridiflora subsp. kuenkeleana     1996      Azerbaydzhan
Epipactis viridiflora        1989    Bosnia-Herzegovina
Epipactis viridiflora        1961    Czech Republic
Epipactis viridiflora     1963 2007200720062006  France
Epipactis viridiflora        1996    Germany
Epipactis viridiflora       196619961970   Switzerland

Species

Below is the list of names accepted by the Kew Monocot Checklist. For the complete list - including invalid names - please click here.

World Checklist for Epipactis. With the permission of the Trustees of the Royal Botanic Gardens, Kew Published on the Internet
Database integration using Royal Botanical Gardens Kew for reference
Hybrid  Not a Hybrid ... Click on the ---- to get more data

Epipactis  -----   Zinn, Cat. Pl. Hort. Gott.: 85 (1757), nom. cons.
   Epipactis africana  -----   Rendle, J. Bot. 33: 252 (1895).
   Epipactis albensis  -----   Nováková & Rydlo, Preslia 50: 162 (1978).
      Epipactis albensis var. albensis  -----  .
      Epipactis albensis var. fibri  -----   (Scappat. & Robatsch) P.Delforge, Naturalistes Belges 78: 246 (1997).
   Epipactis x amigoi  -----   J.M.Lewin, Orchidophile (Asnières) 146: 77 (2001).
   Epipactis atromarginata  -----   Seidenf., Opera Bot. 114: 22 (1992).
   Epipactis atrorubens  -----   (Hoffm.) Besser, Prim. Fl. Galic. 2: 220 (1809).
   Epipactis x barlae  -----   A.Camus in E.G.Camus & A.A.Camus, Iconogr. Orchid. Europe: 486 (1929).
   Epipactis x barreana  -----   B.Baumann & H.Baumann, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 20: 39 (1988).
   Epipactis baumanniorum  -----   Ströhle, J. Eur. Orch. 36: 1044 (2004).
   Epipactis x breinerorum  -----   Batousek, J. Eur. Orch. 29: 645 (1997).
   Epipactis x bruxellensis  -----   P.Delforge, Naturalistes Belges 77(4): 221 (1996).
   Epipactis bugacensis  -----   Robatsch, Ber. Arbeitskreis. Heimische Orchid. 7: 13 (1990).
      Epipactis bugacensis subsp. bugacensis  -----  .
      Epipactis bugacensis subsp. rhodanensis  -----   (Gévaudan & Robatsch) Wucherpf., J. Eur. Orchideen 35: 48 (2003).
   Epipactis x capellonensis  -----   B.Baumann & H.Baumann, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 20: 44 (1988).
   Epipactis x cardonneae  -----   J.M.Lewin, Orchidophile (Asnières) 146: 74 (2001).
   Epipactis condensata  -----   Boiss. ex D.P.Young, Jahresber. Naturwiss. Vereins Wuppertal 23: 106 (1970).
   Epipactis x conquensis  -----   Benito & C.E.Hermos., Estud. Mus. Cienc. Nat. Alava 13: 112 (1998).
   Epipactis dunensis  -----   (T.Stephenson & T.A.Stephenson) Godfery, J. Bot. 64: 68 (1926).
   Epipactis fageticola  -----   (C.E.Hermos.) Devillers-Tersch. & Devillers, Naturalistes Belges 80: 302 (1999).
   Epipactis flaminia  -----   P.R.Savelli & Aless., Webbia 49: 26 (1994).
   Epipactis flava  -----   Seidenf., Dansk Bot. Ark. 32: 116 (1978).
   Epipactis x gerbaudiorum  -----   P.Delforge, Naturalistes Belges 78: 187 (1997).
   Epipactis x gevaudanii  -----   P.Delforge, Naturalistes Belges 78: 179 (1997).
   Epipactis gigantea  -----   Douglas ex Hook., Fl. Bor.-Amer. 2: 202 (1839).
   Epipactis x graberi  -----   A.Camus in E.G.Camus & A.A.Camus, Iconogr. Orchid. Europe: 487 (1929).
   Epipactis greuteri  -----   H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 13: 344 (1981).
   Epipactis guegelii  -----   Robatsch, J. Eur. Orch. 28: 767 (1996 publ. 1997).
   Epipactis helleborine  -----   (L.) Crantz, Stirp. Austr. Fasc., ed. 2, 2(6): 467 (1769).
      Epipactis helleborine subsp. bithynica  -----   (Robatsch) Kreutz, Kompend. Eur. Orchid.: 61 (2004).
      Epipactis helleborine subsp. densifolia  -----   (W.Hahn, Passin & R.Wegener) Kreutz, Eurorchis 17: 105 (2005).
      Epipactis helleborine subsp. helleborine  -----  .
      Epipactis helleborine subsp. latina  -----   W.Rossi & E.Klein, Orchidee (Hamburg) 38: 93 (1987).
      Epipactis helleborine subsp. leutei  -----   (Robatsch) Kreutz, Kompend. Eur. Orchid.: 62 (2004).
      Epipactis helleborine subsp. molochina  -----   (P.Delforge) Kreutz, Kompend. Eur. Orchid.: 62 (2004).
      Epipactis helleborine subsp. neerlandica  -----   (Verm.) Buttler, Willdenowia 16: 115 (1986).
      Epipactis helleborine subsp. orbicularis  -----   (K.Richt.) E.Klein, Phyton (Horn) 37: 74 (1997).
      Epipactis helleborine subsp. schubertiorum  -----   (Bartolo, Pulv. & Robatsch) Kreutz, Kompend. Eur. Orchid.: 62 (2004).
      Epipactis helleborine var. tangutica  -----   (Schltr.) S.C.Chen & G.H.Zhu, Novon 13: 423 (2003).
      Epipactis helleborine subsp. transcaucasica  -----   A.P.Khokhr., Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 96(4): 107 (1991).
      Epipactis helleborine subsp. tremolsii  -----   (Pau) E.Klein, Orchidee (Hamburg) 30: 49 (1979).
   Epipactis heraclea  -----   P.Delforge & Kreutz, Naturalistes Belges 84: 22 (2003).
   Epipactis x heterogama  -----   M.Bayer, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 196 (1986).
   Epipactis humilior  -----   (Tang & F.T.Wang) S.C.Chen & G.H.Zhu, Novon 13: 423 (2003).
   Epipactis kleinii  -----   M.B.Crespo & M.R.Lowe & Piera, Taxon 50: 854 (2001).
   Epipactis lapidocampi  -----   E.Klein & Laminger, Phyton (Horn) 44: 187 (2004).
   Epipactis leptochila  -----   (Godfery) Godfery, J. Bot. 59: 146 (1921).
      Epipactis leptochila subsp. aspromontana  -----   (Bartolo, Pulv. & Robatsch) Kreutz, Kompend. Eur. Orchid.: 63 (2004).
      Epipactis leptochila subsp. futakii  -----   (Mered'a & Potucek) Kreutz, Kompend. Eur. Orchid.: 63 (2004).
      Epipactis leptochila subsp. komoricensis  -----   (Mered'a) Kreutz, Kompend. Eur. Orchid.: 63 (2004).
      Epipactis leptochila subsp. leptochila  -----  .
      Epipactis leptochila subsp. maestrazgona  -----   (P.Delforge & Gévaudan) Kreutz, Kompend. Eur. Orchid.: 63 (2004).
      Epipactis leptochila subsp. naousaensis  -----   (Robatsch) Kreutz, Kompend. Eur. Orchid.: 63 (2004).
      Epipactis leptochila subsp. neglecta  -----   Kümpel, Mitt. Arbeitskreises Beob. Heimischer Orchideen 11: 29 (1982).
   Epipactis liestalensis  -----   A.Camus in E.G.Camus & A.A.Camus, Iconogr. Orchid. Europe: 486 (1929).
   Epipactis lusitanica  -----   D.Tyteca, Orchidophile (Asnières) 84: 218 (1988).
   Epipactis magnibracteata  -----   C.Schweinf., Bot. Mus. Leafl. 5: 39 (1937).
   Epipactis mairei  -----   Schltr., Repert. Spec. Nov. Regni Veg. Beih. 4: 55 (1919).
   Epipactis meridionalis  -----   H.Baumann & R.Lorenz, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 20: 656 (1988).
   Epipactis microphylla  -----   (Ehrh.) Sw., Kongl. Vetensk. Acad. Nya Handl. 21: 232 (1800).
   Epipactis muelleri  -----   Godfery, J. Bot. 59: 106 (1921).
      Epipactis muelleri subsp. cerritae  -----   M.P.Grasso, Orchidee (Hamburg), Suppl. 2: 4 (1994).
      Epipactis muelleri subsp. muelleri  -----  .
   Epipactis x nicolosii  -----   M.P.Grasso & Grillo, Orchidophile (Asnières) 162: 223 (2004).
   Epipactis nordeniorum  -----   Robatsch, Mitt. Abt. Bot. Landesmus. "Joanneum" Graz 20: 32 (1991).
   Epipactis ohwii  -----   Fukuy., Bot. Mag. (Tokyo) 48: 298 (1934).
   Epipactis olympica  -----   Robatsch, Linzer Biol. Beitr. 22: 144 (1990).
   Epipactis palustris  -----   (L.) Crantz, Stirp. Austr. Fasc., ed. 2, 2(6): 463 (1769).
   Epipactis papillosa  -----   Franch. & Sav., Enum. Pl. Jap. 2: 519 (1878).
   Epipactis persica  -----   (Soó) Hausskn. ex Nannf., Bot. Not. 1946: 2 (1946).
   Epipactis phyllanthes  -----   G.E.Sm., Gard. Chron. 12: 660 (1852).
      Epipactis phyllanthes var. olarionensis  -----   P.Delforge, Naturalistes Belges 78: 242 (1997).
      Epipactis phyllanthes var. phyllanthes  -----  .
   Epipactis placentina  -----   Bongiorni & P.Grünanger, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 25: 460 (1993).
   Epipactis pontica  -----   Taubenheim, Orchidee (Hamburg) 26: 68 (1975).
   Epipactis x populetorum  -----   Benito & C.E.Hermos., Estud. Mus. Cienc. Nat. Alava 13: 108 (1998).
   Epipactis provincialis  -----   Aubenas & Robatsch, Orchidophile (Asnières) 122: 109 (1996).
   Epipactis x pupplingensis  -----   K.P.Bell, Jahresber. Naturwiss. Vereins Wuppertal 23: 116 (1970).
   Epipactis purpurata  -----   Sm., Engl. Fl. 4: 41 (1828), nom. cons.
   Epipactis rechingeri  -----   Renz, Orchidee (Hamburg) 24: 253 (1973).
   Epipactis x reinekei  -----   M.Bayer, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 202 (1986).
   Epipactis robatschiana  -----   Bartolo, D'Emerico, Pulv., Terrasi & Stuto, Caryologia 56: 441 (2003).
   Epipactis x robatschii  -----   Gévaudan & P.Delforge, Naturalistes Belges 85: 74 (2004).
   Epipactis royleana  -----   Lindl., Gen. Sp. Orchid. Pl.: 461 (1840).
   Epipactis x schmalhausenii  -----   K.Richt., Pl. Eur. 1: 284 (1890).
      Epipactis x schmalhausenii nothosubsp. fleischmannii  -----   (Heimerl) ined..
      Epipactis x schmalhausenii nothosubsp. schmalhausenii  -----  .
   Epipactis x schulzei  -----   P.Fourn., Fl. Compl. Plaine Franç.: 514 (1928).
   Epipactis x soguksuensis  -----   Kreutz, Eurorchis 9: 71 (1997).
   Epipactis stellifera  -----   Di Antonio & Veya, Candollea 56: 204 (2001).
   Epipactis x stephensonii  -----   Godfery, Monogr. Ic. Br. Nat. Orchid.: 76 (1933).
   Epipactis tallosii  -----   A.Molnár & Robatsch, J. Eur. Orch. 28: 789 (1996 publ. 1997).
   Epipactis tenii  -----   Schltr., Repert. Spec. Nov. Regni Veg. 17: 64 (1921).
   Epipactis thunbergii  -----   A.Gray in M.C.Perry, Exp. Jap. 2: 319 (1856).
   Epipactis troodi  -----   H.Lindb., Årsbok-Vuosik. Soc. Sci. Fenn. 20B(7): 11 (1942).
   Epipactis turcica  -----   Kreutz, Eurorchis 9: 61 (1997).
   Epipactis turcomanica  -----   K.P.Popov & Neshat., Izv. Akad. Nauk Turkmensk. S.S.R., Ser. Biol. Nauk 1982(4): 16 (1982).
   Epipactis ulugurica  -----   Mansf., Notizbl. Bot. Gart. Berlin-Dahlem 11: 1060 (1934).
   Epipactis veratrifolia  -----   Boiss. & Hohen. in P.E.Boissier, Diagn. Pl. Orient. 13: 11 (1854).
   Epipactis x vermionensis  -----   B.Baumann & H.Baumann, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 20: 48 (1988).
   Epipactis xanthophaea  -----   Schltr., Repert. Spec. Nov. Regni Veg. Beih. 12: 341 (1922).

1) - J. Mesjasz-Przybylowicz, W.J. Przybylowicz, B. Godzik, K. Turnau. 37th Microscopy Society of South Africa Conference, Johannesburg, 2 - 4 December 1998. Proceedings - Vol.28, p. 64